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Towers and upgrades in Bloons TD 4

Dart Monkey upgrades	Piercing Dart • Long Range Darts • Spike-o-pult • Triple Shot
Tack Shooter upgrades	Faster Shooting • Extra Range • Blade Shooter • Ring of Fire
Boomerang Thrower upgrades	Multi Target • Sonic Boom • Glaive Thrower • Lightsabre Thrower
Bomb Tower upgrades	Bigger Bombs • Longer Range • Missile Launcher • MOAB Mauler
Ice Tower upgrades	Improved Ice Tower • Permafrost • Snap Freeze • Arctic Wind
Glue Gunner upgrades	Stickier Glue • Glue Soak • Corrosive Glue • Glue Splatter
Monkey Beacon upgrades	Wider Influence • Jungle Drums • Sonar Beacon • Monkey Storm
Super Monkey upgrades	Super Range • Laser Vision • Plasma Vision • Sun God
Mortar Tower upgrades	Greater Accuracy • Rapid Reload • Bloon buster • Mortar Battery
Monkey Ace upgrades	Pineapple Express • Spy Plane • Rapid Fire • Operation: Dart Storm
Monkey Apprentice upgrades	Intense Magic • Fireball • Summon Whirlwind • Tempest Tornado
Banana Farm upgrades	More Bananas • Banana Plantation • Banana Republic • Banana Research Facility
Monkey Buccaneer upgrades	Grape Shot • Crow's Nest • Longer Cannons • Battleship
Dartling Gun upgrades	Steady Barrels • Powerful Shots • Increased Barrel Spin • Laser Cannon
Spike Factory upgrades	Increased Production • Bigger Stacks • White Hot Spikes • Spike Storm
Bee Keeper upgrades	Fast Deploy • Big Hive • Killer Bees • The Swarm
Premium Upgrades	Frag Bombs • Double Ranga • Explosive Darts • Crystal Ball • Dreadnought • Digital Amplification • Monkey Tycoon • Healthy Bananas • Double Cash

List of upgrades