| Dart Monkey
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| Path 1
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Sharp Shots • Razor Sharp Shots • Spike-o-pult • Juggernaut • Ultra-Juggernaut
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| Path 2
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Quick Shots • Very Quick Shots • Triple Shot • Super Monkey Fan Club • Plasma Monkey Fan Club
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| Path 3
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Long Range Darts • Enhanced Eyesight • Crossbow • Sharp Shooter • Crossbow Master
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| Boomerang Monkey
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| Path 1
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Improved Rangs • Glaives • Glaive Ricochet • M.O.A.R Glaives • Glaive Lord
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| Path 2
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Faster Throwing • Faster Rangs • Bionic Boomerang • Turbo Charge • Perma Charge
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| Path 3
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Long Range Rangs • Red Hot Rangs • Kylie Boomerang • MOAB Press • MOAB Domination
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| Bomb Shooter
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| Path 1
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Bigger Bombs • Heavy Bombs • Really Big Bombs • Bloon Impact • Bloon Crush
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| Path 2
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Faster Reload • Missile Launcher • MOAB Mauler • MOAB Assassin • MOAB Eliminator
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| Path 3
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Extra Range • Frag Bombs • Cluster Bombs • Recursive Cluster • Bomb Blitz
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| Tack Shooter
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| Path 1
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Faster Shooting • Even Faster Shooting • Hot Shots • Ring of Fire • Inferno Ring
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| Path 2
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Long Range Tacks • Super Range Tacks • Blade Shooter • Blade Maelstrom • Super Maelstrom
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| Path 3
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More Tacks • Even More Tacks • Tack Sprayer • Overdrive • The Tack Zone
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| Ice Monkey
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| Path 1
|
Permafrost • Cold Snap • Ice Shards • Embrittlement • Super Brittle
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| Path 2
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Enhanced Freeze • Deep Freeze • Arctic Wind • Snowstorm • Absolute Zero
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| Path 3
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Larger Radius • Re-Freeze • Cryo Cannon • Icicles • Icicle Impale
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| Removed
|
Metal Freeze
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| Glue Gunner
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| Path 1
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Glue Soak • Corrosive Glue • Bloon Dissolver • Bloon Liquefier • The Bloon Solver
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| Path 2
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Bigger Globs • Glue Splatter • Glue Hose • Glue Strike • Glue Storm
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| Path 3
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Stickier Glue • Stronger Glue • MOAB Glue • Relentless Glue • Super Glue
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| Sniper Monkey
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| Path 1
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Full Metal Jacket • Large Calibre • Deadly Precision • Maim MOAB • Cripple MOAB
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| Path 2
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Night Vision Goggles • Shrapnel Shot • Bouncing Bullet • Supply Drop • Elite Sniper
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| Path 3
|
Fast Firing • Even Faster Firing • Semi-Automatic • Full Auto Rifle • Elite Defender
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| Monkey Sub
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| Path 1
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Longer Range • Advanced Intel • Submerge and Support • Bloontonium Reactor • Energizer
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| Path 2
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Barbed Darts • Heat-tipped Darts • Ballistic Missile • First Strike Capability • Pre-emptive Strike
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| Path 3
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Twin Guns • Airburst Darts • Triple Guns • Armor Piercing Darts • Sub Commander
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| Monkey Buccaneer
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| Path 1
|
Faster Shooting • Double Shot • Destroyer • Aircraft Carrier • Carrier Flagship
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| Path 2
|
Grape Shot • Hot Shot • Cannon Ship • Monkey Pirates • Pirate Lord
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| Path 3
|
Long Range • Crow's Nest • Merchantman • Favored Trades • Trade Empire
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| Monkey Ace
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| Path 1
|
Rapid Fire • Lots More Darts • Fighter Plane • Operation: Dart Storm • Sky Shredder
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| Path 2
|
Exploding Pineapple • Spy Plane • Bomber Ace • Ground Zero • Tsar Bomba
|
| Path 3
|
Sharper Darts • Advanced Navigation • Neva-Miss Targeting • Spectre • Flying Fortress
|
| Removed
|
Centered Path
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| Heli Pilot
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| Path 1
|
Quad Darts • Pursuit • Razor Rotors • Apache Dartship • Apache Prime
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| Path 2
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Bigger Jets • IFR • Downdraft • Support Chinook • Special Poperations
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| Path 3
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Faster Darts • Faster Firing • MOAB Shove • Comanche Defense • Comanche Commander
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| Mortar Monkey
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| Path 1
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Bigger Blast • Bloon Buster • Shell Shock • The Big One • The Biggest One
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| Path 2
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Faster Reload • Rapid Reload • Heavy Shells • Artillery Battery • Pop and Awe
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| Path 3
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Dynamic Targeting • Burny Stuff • Signal Flare • Shattering Shells • Blooncineration
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| Removed
|
Increased Accuracy
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| Dartling Gunner
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| Path 1
|
Focused Firing • Laser Shock • Laser Cannon • Plasma Accelerator • Ray of Doom
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| Path 2
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Advanced Targeting • Faster Barrel Spin • Hydra Rocket Pods • Rocket Storm • M.A.D
|
| Path 3
|
Faster Swivel • Powerful Darts • Buckshot • Bloon Area Denial System • Bloon Exclusion Zone
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| Wizard Monkey
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| Path 1
|
Guided Magic • Arcane Blast • Arcane Mastery • Arcane Spike • Archmage
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| Path 2
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Fireball • Wall of Fire • Dragon's Breath • Summon Phoenix • Wizard Lord Phoenix
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| Path 3
|
Intense Magic • Monkey Sense • Shimmer • Necromancer • Prince of Darkness
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| Super Monkey
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| Path 1
|
Laser Blasts • Plasma Blasts • Sun Avatar • Sun Temple • True Sun God
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| Path 2
|
Super Range • Epic Range • Robo Monkey • Tech Terror • The Anti-Bloon
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| Path 3
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Knockback • Ultravision • Dark Knight • Dark Champion • Legend of the Night
|
| Ninja Monkey
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| Path 1
|
Ninja Discipline • Sharp Shurikens • Double Shot • Bloonjitsu • Grandmaster Ninja
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| Path 2
|
Distraction • Counter-Espionage • Shinobi Tactics • Bloon Sabotage • Grand Saboteur
|
| Path 3
|
Seeking Shuriken • Caltrops • Flash Bomb • Sticky Bomb • Master Bomber
|
| Alchemist
|
| Path 1
|
Larger Potions • Acidic Mixture Dip • Berserker Brew • Stronger Stimulant • Permanent Brew
|
| Path 2
|
Stronger Acid • Perishing Potions • Unstable Concoction • Transforming Tonic • Total Transformation
|
| Path 3
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Faster Throwing • Acid Pool • Lead to Gold • Rubber to Gold • Bloon Master Alchemist
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| Druid
|
| Path 1
|
Hard Thorns • Heart of Thunder • Druid of the Storm • Ball Lightning • Superstorm
|
| Path 2
|
Thorn Swarm • Heart of Oak • Druid of the Jungle • Jungle's Bounty • Spirit of the Forest
|
| Path 3
|
Druidic Reach • Heart of Vengeance • Druid of Wrath • Poplust • Avatar of Wrath
|
| Monkey Village
|
| Path 1
|
Bigger Radius • Jungle Drums • Primary Training • Primary Mentoring • Primary Expertise
|
| Path 2
|
Grow Blocker • Radar Scanner • Monkey Intelligence Bureau • Call to Arms • Homeland Defense
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| Path 3
|
Monkey Business • Monkey Commerce • Monkeyconomy • Monkey City • Monkeyopolis
|
| Removed
|
Monkey Town
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| Banana Farm
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| Path 1
|
Increased Production • Greater Production • Banana Plantation • Banana Research Facility • Banana Central
|
| Path 2
|
Long Life Bananas • Valuable Bananas • Monkey Bank • IMF loan • Monkey-Nomics
|
| Path 3
|
Quality Soil • Banana Salvage • Marketplace • Central Market • Monkey Wall Street
|
| Removed
|
EZ Collect
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| Spike Factory
|
| Path 1
|
Bigger Stacks • White Hot Spikes • Spiked Balls • Spiked Mines • Super Mines
|
| Path 2
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Faster Production • Even Faster Production • MOAB SHREDR • Spike Storm • Carpet of Spikes
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| Path 3
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Long Reach • Smart Spikes • Long life spikes • Deadly Spikes • Perma-Spike
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| Engineer Monkey
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| Path 1
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Sentry Gun • Faster Engineering • Sprockets • Sentry Expert • Sentry Champion
|
| Path 2
|
Larger Service Area • Deconstruction • Cleansing Foam • Overclock • Ultraboost
|
| Path 3
|
Oversize Nails • Pin • Double Gun • Bloon Trap • XXXL Trap
|
Upgrade cost increased ― $350 → $400
No cross-path (2-0-0)
Long Range Tacks (2-1-0)
Super Range Tacks (2-2-0)
More Tacks (2-0-1)
Even More Tacks (2-0-2)
Portrait
Upgrade icon